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What is the "Big One"?

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Not much of a paper currency collector but curious what "The BIg One" is in the currency world?


It obviously will probably depend on the country but I am curious for a note/bill/certificate that COULD very well exist but none are known, at least, in private hands.


I am thinking comparable to the US 1849 $20 Gold Piece or the 1964-D Peace Dollar.


Edit: Bills/Notes that are speculative to ever of even existed in the first place count.

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